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Here, 15N was applied in the nutrient patch to determine actual nitrogen http://www.selleckchem.com/products/sch772984.html uptake, giving insight in the foraging strategy of the roots in a specific patch. As a null model for nutrient uptake, we hypothesize that nutrient uptake (15N) from a patch is proportional to RLD of a particular species in that patch. The experiment was carried out with species that co-occur in riverine floodplains in the Netherlands. Because it is difficult to separate roots of different species, we used two species that significantly differed in their root morphology, the herb Rumex palustris Sm. and the grass Agrostis stolonifera L. These species are known to display a differential ability to respond to nutrient heterogeneity. R.?palustris displays a high degree of root proliferation in heterogeneous soil (Jansen, Van De Steeg & De Kroon 2005; Jansen et?al. 2006), whereas A.?stolonifera is less responsive (personal observation). R.?palustris is a fast-growing, biennal species from frequently, but not http://www.selleckchem.com/products/Bortezomib.html permanently, flooded soils (Voesenek et?al. 2004). It occurs on species-poor mudflats around stagnant waters in the floodplain. A.?stolonifera is the most characteristic grass species of frequently and prolonged flooded grassland (Van Eck et?al. 2004). This perennial grass species has an elongated stoloniferous growth form and competes for open spaces in the vegetation (Lenssen et?al. 2005). Rumex palustris seeds were collected in 2000 from the Ewijkse Waard, a field site in the floodplains of the river Rhine near Nijmegen http://www.selleck.cn/products/gdc-0068.html in the east of the Netherlands. Seeds of R.?palustris were germinated on wet filter paper in Petri dishes in a growth chamber (16?h light, 25��5?��C; 8?h dark, 10?��C). Rooted shoots of separate clones of A.?stolonifera were collected in the Ewijkse Waard in 2004, at least 50?m apart. These were potted and kept in a greenhouse. Seedlings of R.?palustris and cuttings of A.?stolonifera, consisting of two leaves with roots, were planted on a 1:1 (v:v) mixture of sand and loamy sand in 5��5?cm diameter pots, with an open mesh pot inside. The total plant biomass at the start of the experiment was the same for both species (one-way anova, F1,22?=?0��01; P?>?0��05), although the shoot biomass of A.?stolonifera tended to be higher than that of R.?palustris (mean?��?SE = 0��057?��?0��003?g for R.?palustris, 0��069?��?0��006?g for A.?stolonifera, F1,22?=?3��20; P?