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6.7ensp;Across all species and years, relative growth rate of height increased with K addition and with N and P in combination. Relative growth rate of leaf count trended 8.5% higher with K addition (P?=?0.076). 7.?We also added micronutrients in a parallel experiment. There was no effect http://www.selleckchem.com/products/AZD6244.html of micronutrient addition on any seedling parameter. 8.?Synthesis. K addition affected seedlings by enhancing tissue nutrient concentration, increasing herbivory, reducing root-to-shoot biomass ratio and increasing height growth. Additional effects of N or P on tissue chemistry, herbivory and growth offer support for the multiple limiting resources hypothesis. Our results suggest that seedling growth is limited by nutrients, especially K, even under highly shaded conditions in http://www.selleck.cn/products/Imatinib(STI571).html this lowland tropical forest. Light is considered the most limiting resource in the understorey of tropical forests (Coomes & Grubb 2000), with often http://www.selleckchem.com/products/SP600125.html (K) fertilization experiment and in a parallel micronutrient addition over 8?years to determine the nature of nutrient limitation and identify limiting elements. Several recent meta-analyses of nutrient addition experiments are consistent with multiple limitation of plant growth (Tripler et?al. 2006; Elser et?al. 2007; LeBauer & Treseder 2008). Tropical forests are severely underrepresented in these meta-analyses with stand-level fertilization experiments limited to just a few locations in the lowland tropics (Mirmanto et?al. 1999; Newbery et?al. 2002; Campo & V��zquez-Yanes 2004; Davidson et?al. 2004; Lu et?al. 2010; Wright et?al. 2011).