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These regression equations were used to calculate the amount of coral tissue removed per day by each coral-feeding butterflyfish observed on each transect to provide an overall estimate of the total amount of coral tissue removed by all coral-feeding butterflyfishes per day per 200?m2 at each of the three study sites (Table?1, step 4). Two butterflyfish species (Chaetodon rainfordi and C.?trifascialis) were detected on these transects for which size-specific regression equations are not available. In these cases, regression equations of the closest related species were used (C. aureofasciatus and C.?baronessa respectively) (Bellwood et?al. 2010). The total amount of coral tissue removed per day by coral-feeding butterflyfishes at each site (Table?1, step 4) was then multiplied by the proportion of bites taken by each species on tabular acroporids (A.?hyacinthus and A.?cytherea) http://www.selleckchem.com/products/3-methyladenine.html at each of the three sites based on our feeding observation data (Table?1, step 5) http://www.selleckchem.com/products/PD-0325901.html to determine the total amount of tissue removed from tabular acroporids per 200?m2 per day (Table?1, step 6). Chaetodon aureofasciatus, C.?rainfordi and Chaetodon plebeius are generally rare on the reef crest habitat and accounted for http://www.selleck.cn/products/Cisplatin.html the total amount of coral tissue consumed per day to an equivalent three-dimensional surface area (cm2) of coral (Table?1, step 7). To calculate the proportion of tabular acroporid tissue consumed annually by corallivorous butterflyfishes, we multiplied the areal equivalents of daily coral consumption estimates by 365 (Table?1, step 8) and divided by the actual three-dimensional surface area of tabular acroporids tissue that was available at each site (Table?1, step 9). To calculate the error around our estimate of annual consumption, we repeated these calculations three times; once using the mean value for each variable in all calculations, once using the mean plus one standard error in all calculations and once using the mean minus one standard error, with our final error values representing the deviation from the mean in our three calculations. We do not expect the extrapolation from daily to yearly consumption estimates to be distorted by seasonal differences in feeding rates. Unlike herbivorous fishes, corallivorous butterflyfishes are surprisingly consistent, and no study has documented a significant effect of season on the feeding rates of these fishes (e.g. Cox 1986; Irons 1989; Pratchett 2005 etc.).