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We calculated nested temperature using the method of Rodr��guez-Giron��s & Santamar��a (2006), which provides a measure T of the degree to which species assemblages are nested across sites varying in species richness, with low values of T indicating a high degree of nestedness (Atmar & Patterson 1993; Ulrich, Almeida-Neto & Gotelli 2009). We also generated null model communities to assess the non-randomness of nestedness patterns (Mikl��s & Podani 2004). A meta-community is perfectly nested if species-poor microsites (quadrats) are subsets of progressively more species-rich microsites, and rare or uncommon species are present only in species-rich microsites (Ulrich, Almeida-Neto & Gotelli 2009). In general, rare and uncommon species http://www.selleck.cn/products/ve-821.html are at greater risk of local extinction due to their small population sizes (e.g. Pimm et?al. 1995), and non-random, nested patterns of species occurrences along diversity gradients result from some species being more susceptible to environmental stressors than others (e.g. Bracken & Nielsen 2004). Therefore, a highly nested distribution pattern may indicate a predictable order of species loss from more to less species-rich microsites (Zavaleta & Hulvey 2004). Nested subset analysis can thus be used as an unbiased, realistic estimate of the order of species loss with no a priori assumptions about the ultimate drivers of species loss. Plant species assemblages at Coyote Ridge were most nested in 1991 (T?=?17.86, P?=?0.01), and least nested in 1996 (T?=?38.94, http://www.selleckchem.com/products/Vorinostat-saha.html P? http://www.selleckchem.com/products/Cisplatin.html slopes model (PRISM; http://prism.oregonstate.edu) to estimate annual precipitation for the years 1960�C2009 at Coyote Ridge (Fig.?S1). Although annual precipitation in 1991 was only ?16% below the 50-year average of 567?mm, it immediately followed the most prolonged drought over the past 50?years (1988�C90), in which precipitation was >40% below the 50-year mean for each year (Fig.?S1). This suggests the high degree of nestedness in 1991 was driven by prolonged drought in preceding years. Notably, the 2?years with the next highest degree of nestedness after 1991 were 2007 (T?=?21.84, P?=?0.02) and 2008 (T?=?24.72, P?=?0.01); 2007 had the lowest annual precipitation in the 50-year period examined (262?mm), and 2008 was also a below-average precipitation year (405?mm, ?30% below the 50-year mean). The year with the lowest degree of nestedness, 1996 (T?=?38.94, P?